Oto, Japan’s oldest lion, died Monday of natural causes at the Toyohashi Zoo & Botanical Park in Aichi Prefecture, according to the zoo. She was 25½ years old — equivalent to about 100 human ...

Japan’s oldest lion, Oto, died on Monday at the age of 25, which is equivalent to a century in human years. Born in 1999 in Hokkaido’s Asahiyama Zoo, Oto had lived in the Toyohashi Zoo and ...